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Roman Grain Pests in Britain: Implications for Grain Supply and Agricultural Production

Published online by Cambridge University Press:  19 April 2011

David Smith
Affiliation:
Institute for Archaeology and Antiquity, The University of Birminghamd.n.smith@bham.ac.uk
Harry Kenward
Affiliation:
Department of Archaeology, The University of Yorkhkk1@york.ac.uk

Abstract

It is over 30 years since Paul Buckland first presented a series of arguments concerning beetle (Coleoptera) grain pests: their origin, the timing of their introduction to Britain, and their implications for agricultural production during the Roman occupation. Here we return to the topic in the light of new data from a range of archaeological deposits, including civilian and military sites dating from the earliest period of Roman occupation. Infestation rates and, potentially, grain loss may have been high throughout Roman Britain, though many infestations may have been in equine feed. Beetle grain pests are not recorded in Britain prior to the Roman invasion, and it appears that they were absent, or extremely rare, in the early medieval period and up to the Norman Conquest. This pattern of occurrence is reviewed and it is suggested that ecological theory offers an explanation which is in accord with supposed socio-economic changes and trade. The role of grain pests is considered in the economic modelling of Romano-British agriculture.

Type
Articles
Copyright
Copyright © The Author(s) 2011. Published by The Society for the Promotion of Roman Studies

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References

BIBLIOGRAPHY104

Aitken, A.D. 1975: Insect Travellers. Volume 1. Coleoptera, LondonGoogle Scholar
Allison, E.P., Hall, A.R., Kenward, H.K., and O'Connor, T.P. 1990: Environmental Evidence from the Roman Settlement, Staniwells Farm, Hibaldstow, Humberside, 1988, Reports from the Environmental Archaeology Unit, York 1990/05, YorkGoogle Scholar
Allison, E.P., Hutchinson, A., Jones, A.K.G., Kenward, H.K., and Morgan, L.M. 1991: passim in McCarthy 1991, 849Google Scholar
Armstrong, M.T., and Howe, R.W. 1963: ‘The saw-toothed grain beetle (Oryzaephilus surinamensis) in home grown grain’, Journal of Agricultural Engineering Research 8, 256–61Google Scholar
Barber, K.E., Maddy, D., Rose, N., Stevenson, A.C., Stoneman, R., and Thompson, R. 2000: ‘Replicated proxy-climate signals over the last 2000 yr from two distant UK peat bogs: new evidence for regional palaeoclimate teleconnections’, Quaternary Science Reviews 19, 481–7CrossRefGoogle Scholar
Beavis, I.C. 1988: Insects and Other Invertebrates in Classical Antiquity, ExeterCrossRefGoogle Scholar
Bergh, J.-E., Hansen, L.S., Jensen, K.-M.V., and Nielsen, P.V. 2003: ‘The effect of anoxic treatment on the larvae of six species of dermestids (Coleoptera)’, Journal of Applied Entomology 127, 317–21CrossRefGoogle Scholar
Bidwell, P., and Speak, S. 1994: Excavations at South Shields Roman Fort I, Society of Antiquaries of Newcastle upon Tyne with Tyne and Wear Museums Monograph 4, Newcastle upon TyneGoogle Scholar
Briffa, K.R. 2000: ‘Annual climate variability in the Holocene: interpreting the message of ancient trees’, Quaternary Science Reviews 19, 87105CrossRefGoogle Scholar
Brunt, P.A. 1971: Italian Manpower 225 BC–AD 14, OxfordGoogle Scholar
Büchner, S., and Wolf, G. 1997: ‘Der Kornkäfer – Sitophilus granarius (Linné) – aus einer bandkeramischen Grube bei Göttingen’, Archäologisches Korrespondenzblatt 27, 211–20Google Scholar
Buckland, P.C. 1976: The Environmental Evidence from the Church Street Roman Sewer System, The Archaeology of York 14.1, LondonGoogle Scholar
Buckland, P.C. 1978: ‘Cereal production, storage and population: a caveat’, in Limbrey and Evans 1978, 43–5Google Scholar
Buckland, P.C. 1981: ‘The early dispersal of insect pests of stored products as indicated by archaeological records’, Journal of Stored Products Research 17, 112CrossRefGoogle Scholar
Buckland, P.C. 1982: ‘The Malton burnt grain: a cautionary tale’, Yorkshire Archaeological Journal 54, 5361Google Scholar
Buckland, P.C. 1996: ‘Insects’, in J. May, Dragonby. Report on Excavations at an Iron Age and Romano-British Settlement in North Lincolnshire, Oxbow Monograph 61, Oxford, 165–70Google Scholar
Buckland, P.I., and Buckland, P.C. 2006: Bugs Coleopteran Ecology Package (Versions: BugsCEP v7.61; Bugsdata v7.09; BugsMCR v2.0; BugStats v1.2) [Downloaded December 2006] www.bugscep.comGoogle Scholar
Buxton, K., and Howard-Davis, C. 2000: Bremetenacum. Excavations at Roman Ribchester 1980, 1989–1990, Lancaster Imprints Series 9, LancasterGoogle Scholar
Campbell, G. 2000: ‘Plant utilization: the evidence from charred plant remains’, in Cunliffe 2000, 4559Google Scholar
Candura, G.S. 1932: ‘Contributo alla conoscenza morfologica e biologica dello struggigrano (Tenebroides mauritanicus L.)’, Bolletino Laboratorio di Zoologia Generale e Agraria Portici 27, 156Google Scholar
Carrott, J., Issitt, M., Kenward, H., Large, F., McKenna, B., and Skidmore, P. 1995: Insect and Other Invertebrate Remains from Excavations at Four Sites in Lincoln (site codes: WN87, WNW88, WF89 and WO89): Technical Report, Reports from the Environmental Archaeology Unit, York 95/10, YorkGoogle Scholar
Carruthers, W. 1993: A Charred Grain Deposit from Ambleside Roman Fort, Cumbria, Ancient Monuments Laboratory Report 11/93, LondonGoogle Scholar
Chiverell, R.C. 2001: ‘A proxy record of late Holocene climate change from May Moss, northeast England’, Journal of Quaternary Science 16, 9293.0.CO;2-K>CrossRefGoogle Scholar
Coombs, C.W., and Freeman, J.A. 1955: ‘The insect fauna of an empty granary’, Journal of Entomological Research 46, 399417Google Scholar
Coombs, C.W., and Woodroffe, G.E. 1963: ‘An experimental demonstration of ecological succession in an insect population breeding in stored wheat’, Journal of Animal Ecology 32, 271–9CrossRefGoogle Scholar
Coope, G.R., and Osborne, P.J. 1968: ‘Report on the Coleopterous fauna of the Roman well at Barnsley Park, Gloucestershire’, Transactions of the Bristol and Gloucestershire Archaeological Society 86, 84–7Google Scholar
Cowie, R. 2000: ‘Londinium to Lundenwic: early and middle Saxon archaeology in the London region’, in Haynes, I., Sheldon, H. and Hannigan, L. (eds), London Underground: The Archaeology of a City, Oxford, 175205CrossRefGoogle Scholar
Cowie, R., and Harding, C. 2000: ‘Saxon settlement and economy from the Dark Ages to Domesday’, in The Archaeology of Greater London: an Assessment of Archaeological Evidence for Human Presence in the Area Now Covered by Greater London, Museum of London Archaeological Service, London, 171–98Google Scholar
Cunliffe, B. 1992: ‘Pits, preconceptions and propitiation in the British Iron Age’, Oxford Journal of Archaeology 11, 6983CrossRefGoogle Scholar
Cunliffe, B. 2000: The Danebury Environs Programme. The Prehistory of a Wessex Landscape, Vol. 1: Introduction, English Heritage and Oxford University Committee of Archaeology Monograph 48, LondonGoogle Scholar
De Angelis, F. 2000: ‘Estimating the agricultural base of Greek Sicily’, Papers of the British School at Rome 68, 111–48CrossRefGoogle Scholar
de Moulins, D. 1990: ‘Environmental analysis’, in Maloney, C., The Upper Walbrook Valley in the Roman Period, Council for British Archaeology Research Report 69, London, 85115Google Scholar
Dendy, A., and Elkington, H.D. 1920: Report on the Effect of Air-tight Storage upon Grain Insects, Grain Pests Committee Report 6, LondonGoogle Scholar
Dobney, K., Kenward, H., Ottaway, P., and Donel, L. 1998: ‘Down, but not out: biological evidence for complex economic organisation in Lincoln in the late fourth century’, Antiquity 72, 417–24CrossRefGoogle Scholar
Elias, S.A. 2010: Advances in Quaternary Entomology, Developments in Quaternary Sciences 12, AmsterdamGoogle Scholar
Freeman, P. 1980: Common Insect Pests of Stored Products, LondonGoogle Scholar
Garnsey, P. 1988: Famine and Food Supply in the Graeco-Roman World, CambridgeCrossRefGoogle Scholar
Hall, A.R., and Kenward, H.K. 1990: Environmental Evidence from the Colonia: General Accident and Rougier Street, The Archaeology of York 14.6, LondonGoogle Scholar
Hall, A.R., and Kenward, H.K. 2006: ‘Development-driven archaeology: bane or boon for bioarchaeology?’, Oxford Journal of Archaeology 25, 213–24CrossRefGoogle Scholar
Hall, A., and Kenward, H. forthcoming: ‘Dung and stable manure on waterlogged archaeological occupation sites: some ruminations on the evidence from plant and invertebrate remains’, in Jones, R. (ed.), Manure Matters: Archaeological, Historical and Ethnographic Perspectives, FarnhamGoogle Scholar
Hall, A.R., Kenward, H.K., and Williams, D. 1980: Environmental Evidence from Roman Deposits in Skeldergate, The Archaeology of York 14.3, LondonGoogle Scholar
Hall, R.A., and Kenward, H.K. 1976: ‘Biological evidence for the usage of Roman riverside warehouses at York’, Britannia 7, 274–6CrossRefGoogle Scholar
Halstead, P., and O'Shea, J. 1989: Bad Year Economics, CambridgeCrossRefGoogle Scholar
Hamilton, J. 2000: ‘Animal husbandry: the evidence from the animal bone’, in Cunliffe 2000, 5976Google Scholar
Haughton, J.T. 2004: Global Warming: The Complete Briefing (3rd edn), CambridgeCrossRefGoogle Scholar
Hill, J.D. 1995: Ritual and Rubbish in the Iron Age of Wessex, BAR British Series 242, OxfordGoogle Scholar
Hunter, F.A., Tulloch, B.M., and Lamborne, M.G. 1973: ‘Insects and mites of maltings in the East Midlands of England’, Journal of Stored Products Research 9, 119–41CrossRefGoogle Scholar
Hyde, M.B. 1962: ‘Airtight storage of grain’, Annals of Applied Biology 50, 362–4CrossRefGoogle Scholar
Hyde, M.B., and Oxley, T.A. 1960: ‘Experiments on the airtight storage of damp grain. 1. Introduction, effect on the grain and the intergranular atmosphere’, Annals of Applied Biology 48, 687710CrossRefGoogle Scholar
Jones, M.K. 1985: ‘Archaeobotany beyond subsistence reconstruction’, in Barker, G. and Gamble, C. (eds), Beyond Domestication in Prehistoric Europe, London, 107–28Google Scholar
Jongman, W. 1991: The Economy and Society of Pompeii, AmsterdamGoogle Scholar
Jongman, W. 2003: ‘Slavery and the growth of Rome: the transformation of Italy in the second and first centuries BCE’, in Wolfe, G. (ed.), Rome the Cosmopolis, Cambridge, 100–22Google Scholar
Kenward, H.K. 1997: ‘Synanthropic insects and the size, remoteness and longevity of archaeological occupation sites: applying concepts from biogeography to past “islands” of human occupation’, Quaternary Proceedings 5, 135–52Google Scholar
Kenward, H.K. 2004: ‘Do insect remains from historic-period archaeological occupation sites track climate change in Northern England?’, Environmental Archaeology 9, 4759CrossRefGoogle Scholar
Kenward, H.K. 2009: Invertebrates in Archaeology in the North of England, English Heritage Research Department Report Series 12/2009, http://research.english-heritage.org.uk/report/?14728Google Scholar
Kenward, H., and Carrott, J. 2006: ‘Insect species associations characterise past occupation sites’, Journal of Archaeological Science 33, 1452–73CrossRefGoogle Scholar
Kenward, H.K., and Hall, A.R. 1995: Biological Evidence from Anglo-Scandinavian Deposits at 16–22 Coppergate, The Archaeology of York 14.7, YorkGoogle Scholar
Kenward, H.K., and Hall, A.R. 1997: ‘Enhancing bioarchaeological interpretation using indicator groups: stable manure as a paradigm’, Journal of Archaeological Science 24, 663–73.CrossRefGoogle Scholar
Kenward, H.K., and Hall, A.R. 2006: ‘Easily decayed organic remains in urban archaeological deposits: value, threats, research directions and conservation’, in Brinkkemper, O., Deeben, J., van Doesburg, J., Hallewas, D., Theunissen, E.M. and Verlinde, A.D. (eds), Vakken in vlakken. Archeologische kennis in lagen, Nederlandse Archeologische Rapporten 32, Amersfoort, 183–98Google Scholar
Kenward, H.K., and Whitehouse, N.J. 2010: ‘Insects’, in O'Connor, T. and Sykes, N. (eds), Extinctions and Invasions: A Social History of British Fauna, Oxford, 181–9 and 197–231 (bibliography)CrossRefGoogle Scholar
Kenward, H.K., and Williams, D. 1979: Biological Evidence from the Roman Warehouses in Coney Street, The Archaeology of York 14.2, LondonGoogle Scholar
Kenward, H.K., Allison, E.P., Dainton, M., Kemenes, I.K., and Carrott, J.B. 1992: Evidence from Insect Remains and Parasite Eggs from Old Grapes Lane A, The Lanes, Carlisle: Technical Report, Ancient Monuments Laboratory Report 78/92, LondonGoogle Scholar
Kenward, H.K., Allison, E.P., Morgan, L.M., Jones, A.K.G., and Hutchinson, A.R. 1991: ‘The insect and parasite remains’, in McCarthy 1991, 6572Google Scholar
Kenward, H.K., Hall, A.R., and Jones, A.K.G. 1980: ‘A tested set of techniques for the extraction of plant and animal macrofossils from waterlogged archaeological deposits’, Science and Archaeology 22, 315Google Scholar
Kenward, H.K., Hall, A.R., and Jones, A.K.G. 1986: Environmental Evidence from a Roman Well and Anglian Pits in the Legionary Fortress, The Archaeology of York 14.5, LondonGoogle Scholar
Kenward, H., King, G., and Hall, A. 2008: ‘Carbonised insects: rare, overlooked or destroyed by sample processing?’, in Abstracts: Association for Environmental Archaeology, The Consequences of Fire. AEA 2008 Annual Conference, 12.–14. September 2008, Århus, Denmar, Moesgård Museum, Højbjerg, 33–5 http://www.moesmus.dk/archive/pdf_dokumenter/pdf%20konservering%20og%20naturvidenskab/AEA%202008/AEA%202008%20the%20consequences%20of%20fire.pdfGoogle Scholar
Kislev, M.E., and Melamed, Y. 2000: ‘Ancient infested wheat and horsebean from Horbat Rosh Zayit’, in Gal, A. and Alexandre, Y. (eds), Horbat Rosh Zayit. An Iron Age Storage Fort and Village, Jerusalem, 206–20CrossRefGoogle Scholar
Koch, K. 1970: ‘Subfossile Kaferreste aus romerzeitlichen und mittelalterlichen Ausgrabungen im Rheinland’, Entomologische Blatter 66, 4156Google Scholar
Koch, K. 1971: ‘Zur Untersuchung subfossiler Käferreste aus römerzeitlichen und mittelalterlichen Ausgrabungen im Rheinland’, Rheinische Ausgrabungen 10, 373448 + pls 16–27Google Scholar
Langdon, P.G., Barber, K.E., and Hughes, P.D.M. 2003: ‘A 7500-year peat-based palaeoclimatic reconstruction and evidence for an 1100-year cyclicity in bog surface wetness from Temple Hill Moss, Pentland Hills, southeast Scotland’, Quaternary Science Reviews 22, 259–74CrossRefGoogle Scholar
Langdon, P.G., Barber, K.E., and Lomas-Clarke, S.E. 2004: ‘Reconstructing climate and environmental change in northern England through chironomid and pollen analyses: evidence from Talkin Tarn, Cumbria’, Journal of Paleolimnology 32, 197213CrossRefGoogle Scholar
Large, F., Kenward, H., Carrott, J., Nicholson, C., and Kent, P. 1994: Insect and Other Invertebrate Remains from the Roman Fort at Ribchester, Lancashire (site code RB89): Technical Report, Reports from the Environmental Archaeology Unit, York 94/11, YorkGoogle Scholar
Limbrey, S., and Evans, J.G. (eds) 1978: The Effect of Man on the Landscape: the Lowland Zone, Council for British Archaeology Research Report 21, LondonGoogle Scholar
Longstaff, B.C. 1981: ‘Biology of the grain pest species of the genus Sitophilus (Coleoptera: Curculionidae): a critical review’, Protection Ecology 2, 83130Google Scholar
Mason, L., and Strait, C.A. 1998: ‘Stored product integrated pest management with extreme temperatures’, in Hallman, G.J. and Denlinger, D.L. (eds), Temperature Sensitivity in Insects and Application in Integrated Pest Management, Boulder, 141–77Google Scholar
Mauquoy, D., and Barber, K. 1999: ‘A replicated 3000 yr proxy-climate record from Coom Rigg Moss and Felecia Moss, the Borders Mires, northern England’, Journal of Quaternary Science 14, 263–753.0.CO;2-W>CrossRefGoogle Scholar
McCarthy, M.R. 1991: The Roman Waterlogged Remains and Later Features at Castle Street, Carlisle: Excavations 1981–2: Fascicule 1, The Stratigraphic Sequence and the Environmental Remains, Cumberland and Westmorland Antiquarian and Archaeological Society Research Series 5.1, Kendal (microfiche)Google Scholar
McDermott, F., Mattey, D.P., and Hawkesworth, C. 2001: ‘Centennial-scale Holocene climate variability revealed by a high-resolution speleothem δ18O record from SW Ireland’, Science 294, 1328–31CrossRefGoogle ScholarPubMed
McFarlane, J.A. 1989: Guidelines for Pest Management Research to Reduce Stored Food Losses Caused by Insects and Mites, Overseas Development and Natural Resources Institute Bulletin 22, Chatham, KentGoogle Scholar
Morris, M., and Smith, D.N. 2007: ‘Insect remains’, in Bowsher, D., Dyson, T., Holder, N. and Howell, I., The London Guildhall; an Archaeological History of a Neighbourhood from Early Mediveal to Modern Times, Museum of London Archaeology Service Monograph 36, London, 486–7Google Scholar
Munro, J.W. 1966: Pests of Stored Products, LondonGoogle Scholar
Osborne, P.J. 1971a: ‘The insect fauna from the Roman harbour’, in Cunliffe, B.W. (ed.), Excavations at Fishbourne, 1961–1969, Reports of the Research Committee of the Society of Antiquaries of London 27, Oxford, 393–9Google Scholar
Osborne, P.J. 1971b: ‘An insect fauna from the Roman site at Alcester, Warwickshire’, Britannia 2, 156–65CrossRefGoogle Scholar
Osborne, P.J. 1977: ‘Stored product beetles from a Roman site at Droitwich, England’, Journal of Stored Products Research 13, 203–4CrossRefGoogle Scholar
Osborne, P.J. 1994: ‘The insects’, in Bidwell and Speak 1994, 266Google Scholar
Oxley, T.A. 1948: The Scientific Principles of Grain Storage, LiverpoolGoogle Scholar
Pals, J.P., and Hakbijl, T. 1992: ‘Weed and insect infestation of a grain cargo in a ship at the Roman fort of Laurium in Woerden (Province of Zuid-Holland)’, Review of Palaeobotany and Palynology 73, 287300CrossRefGoogle Scholar
Panagiotakopulu, E. 2000: Archaeology and Entomology in the Eastern Mediterranean: Research into the History of Insect Synanthropy in Greece and Egypt, BAR International Series S836, OxfordCrossRefGoogle Scholar
Panagiotakopulu, E. 2001: ‘New records for ancient pests: Archaeoentomology in Egypt’, Journal of Archaeological Science 28, 1235–46CrossRefGoogle Scholar
Panagiotakopulu, E., and Buckland, P.C. 1991: ‘Insect pests of stored products from Late Bronze Age Santorini, Greece’, Journal of Stored Products Research 27, 179–84CrossRefGoogle Scholar
Payne, T.S. 2002: ‘Harvest and storage management of wheat’, in Curtis, B.C.Rajaram, S. and Macpherson, H.G. (eds), Bread Wheat Improvement and Production, FAO Plant Production and Protection Series 30, Rome http://www.fao.org/DOCREP/006/Y4011E/y4011e0u.htm#bm30 (accessed 20 March 2009)Google Scholar
Reynolds, P.J. 1974: ‘Experimental Iron Age storage pits: an interim report’, Proceedings of the Prehistoric Society 40, 118–31CrossRefGoogle Scholar
Robertson, A., Tomlinson, P., and Kenward, H.K. 1989: Plant and Insect Remains from Coffee Yard, York, Reports from the Environmental Archaeology Unit, York 1989/12Google Scholar
Robinson, M.A. 1978: ‘A comparison between the effects of man on the environment of the first gravel terrace and flood-plain of the Upper Thames Valley during the Iron Age and Roman periods’, in Limbrey and Evans 1978, 3543Google Scholar
Robinson, M.A. 1979: ‘The biological evidence’, in Lambrick, G. and Robinson, M.A., Iron Age and Roman Riverside Settlements at Farmoor, Oxfordshire, Council for British Archaeology Research Report 32, London, 77133Google Scholar
Robinson, M.A. 1991: ‘The Neolithic and Late Bronze Age insect assemblages’, in Needham, S. (ed.), Excavation and Salvage at Runnymede Bridge, 1978: the Late Bronze Age Waterfront Site, London, 277325Google Scholar
Robinson, M.A. 1993: ‘The Iron Age environmental evidence’, in Allen, T.G. and Robinson, M.A. (eds), The Prehistoric Landscape and Iron Age Enclosed Settlement at Mingies Ditch, Hardwick-with-Yelford, Oxon, Thames Valley Landscapes: The Windrush Valley Volume 2, Oxford, 101–20Google Scholar
Robinson, M.A. 2007: ‘The environmental archaeology of the Cotswold Water Park’, in Miles, D., Palmer, S., Smith, A. and Jones, G., Iron Age and Roman Settlement in the Upper Thames Valley: Excavations at Claydon Pike and Other Sites within the Cotswold Water Park, Thames Valley Landscapes Monograph 26, Oxford, 355–63Google Scholar
Rosenstein, N. 2004: Rome at War. Farms, Families, and Death in the Middle Republic, LondonGoogle Scholar
Rowsome, P. 2000: Heart of the City: Roman, Medieval and Modern London Revealed by Archaeology at 1 Poultry, LondonGoogle Scholar
Salmond, K.F. 1957: ‘The insect and mite fauna of a Scottish flour mill’, Bulletin of Entomological Research 47, 621–30CrossRefGoogle Scholar
Schmidt, E. 1998: ‘Der Kornkäfer Sitophilus granarius Schoen. Curculionidae aus der Schuttschicht des bandkeramischen Brunnens von Erkelenz-Kückhoven’, in Koschik, H. (ed.), Brunnen der Jungsteinzeit, Internationales Symposium Erkelenz, 27–29 Oktober 1997, Materialien zur Bodendenkmalpflege im Rheinland 11, Köln, 261–9Google Scholar
Sigaut, F. 1988: ‘A method for identifying grain storage techniques and its application for European agricultural history’, Tools and Tillage 6, 332Google Scholar
Smith, D.N. 1997: ‘The insect fauna’, in Thomas, C., Sloane, B. and Philpotts, C. (eds), Excavations at the Priory and Hospital of St Mary Spital, London, Museum of London Archaeology Service Monograph 1, London, 245–7Google Scholar
Smith, D.N. 2004. ‘The insect remains from the well’, in Bishop, M.C., Inveresk Gate: Excavations in the Roman Civil Settlement at Inveresk, East Lothian, 1996–2000, STAR Monograph 7, Loanhead, 81–8Google Scholar
Smith, D.N. 2010: ‘The insect remains’, in Howard-Davis, C., The Carlisle Millennium Project: Excavations in Carlisle 1998–2001. Volume 2: The Finds, Lancaster Imprints 15, Lancaster, 921–5Google Scholar
Smith, D.N. 2011: ‘The insect remains’, in Hill, J. and Rowsome, P., Roman London and the Walbrook Stream Crossing: Excavations at 1 Poultry and Vicinity, City of London, Museum of London Archaeology Service Monograph 37, London, 559–63Google Scholar
Smith, D.N. forthcoming a: ‘The insects from Northfleet’, in Andrews, P., Biddulph, E., Hardy, A. and Smith, A., Settling the Ebbsfleet Valley: CTRL Excavations at Springhead and Northfleet, Kent. The Late Iron Age, Roman, Saxon and Medieval Landscape, Oxford/Wessex ArchaeologyGoogle Scholar
Smith, D.N. forthcoming b: ‘The insect remains’, in Burch, M., Treveil, P. and Keene, D., The Development of Early Medieval and Later Poultry and Cheapside: Excavations at 1 Poultry and Vicinity, City of London, Museum of London Archaeology Service Monograph, LondonGoogle Scholar
Smith, D.N., and Chandler, G. 2004: ‘Insect remains’, in Sloane, B. and Malcolm, G. (eds), Excavations at the Priory of the Order of the Hospital of St John of Jerusalem, Clerkenwell, London, Museum of London Archaeology Service Monograph 20, London, 389–94Google Scholar
Smith, D.N., and Morris, M. 2008: ‘Insect remains’, in Bateman, N., Cowan, C. and Wroe-Brown, R., London's Roman Amphitheatre: Guildhall Yard, City of London, Museum of London Archaeology Service Monograph 35, London, 218–19Google Scholar
Smith, D.N., and Tetlow, E. 2004: The Insect Remains from Gresham Street, University of Birmingham Environmental Archaeology Services Report 85, unpub. specialist reportGoogle Scholar
Solomon, M.E., and Adamson, B.E. 1956: ‘The powers of survival of storage and domestic pests under winter conditions in Britain’, Bulletin of Entomological Research 46, 311–55CrossRefGoogle Scholar
Thomas, E.L., and Shepard, H.H. 1940: ‘The influence of temperature, moisture, and food upon the development and survival of the saw-toothed grain beetle’, Journal of Agricultural Research 60, 605–15Google Scholar
Tyler, P.S., and Boxall, R.A. 1984: ‘Post-harvest loss reduction programmes: a decade of activities: what consequences?’, Tropical Stored Product Information 23, 1328Google Scholar
Van der Veen, M. 1988: ‘Carbonised grain from a Roman granary at South Shields, north-east England’, in Kuster, H., Der prähistorische Mensch und seine Umwelt, Festschrift für Udelgard Körber-Grohne zum 65, Stuttgart, 353–65Google Scholar
Van der Veen, M. 1994: ‘Grain from the forecourt granary’, in Bidwell and Speak 1994, 243–58Google Scholar
Van der Veen, M., and Jones, G. 2006: ‘A re-analysis of agricultural production and consumption: implications for understanding the British Iron Age’, Vegetation History and Archaeobotany 15, 217–28CrossRefGoogle Scholar
White, K.D. 1970: Roman Farming, LondonGoogle Scholar
Yvinec, H.-L. 1997: ‘Infestation par les insectes d'un grenier carbonisé de la fin du IIe siècle A.D. à Amiens (Somme)’, L'Entomologiste (Paris) 53, 113–28Google Scholar